Frullania bolanderi

Shoot anisophyllous Capsule valves vertical
Leaf bases not decurrent
Leaves closely imbricate
Perigynium absent
Lobes asymmetrical
Shoots mainly prostrate
Leaf margin entire
Medium: 1.5 - 2.5 mm wide
Rhizoids prominent
Leaves secund
Leaves spreading
Dioicous
Perianth present
Oil-bodies present
Oil-bodies ave 2-12/cell
Oil-bodies colorless
Lobe tips acute
Perianth long emergent
Perianth mouth narrow
Shoots mainly ascending
Lobes folded together
Sinus acute
Oil-bodies granulose
Small: 0.5 - 1.5 mm wide
Leaves divided >0.5
Lobe tips rounded
Lobes rotund
Cell walls unevenly thick
Tinged with bright red
Underleaves present
Underleaves bilobed
Rhizoids tufted
Shoot tips unreduced
Shoot tips attenuate
Microphyllous branches
Shoots pinnately branched
Dark, olive green
Green to reddish brown
Dark brown to blackish
Leaves incubous
Leaves nearly horizontal
Dorsal lobe larger
Ventral lobule flask-like
Trigones concave
Perianth flattened
Perianth beaked
Caducous leaves present
Elaters unispiral